! 4C Species Parameter File ! contains the following data: ! with * marked parameters are sensitive ! ! nspecies number of all species ! nspec_tree number of tree species ! name species name ! Amax maximum age (years) ! yrec stress recovery time (years) ! stol shade tolerance (1-5) ! pfext extinction coefficient ! sigman* root activity rate ! respcoeff respiration coeff. = fraction of gross production respired by the plant ! prg* growth respiration parameter ! prms* maintenance respiration parameters: sapwood, ! prmr fine roots; ! psf* senescence parameters: foliage, ! pss* sapwood, ! psr* fine roots; ! pcnr* N/C ratio of biomass [kg N/ kg C] ! ncon_fol N concentration of folmg / g TM nicht / g C!!? ! ncon_frt N concentration of fine roots [mg/g] ! ncon_crt N concentration of coarse roots [mg/g] ! ncon_tbc N concentration of twigs and branches [mg/g] ! ncon_stem N concentration of stemwood [mg/g] ! reallo_fol reallocation parameter of foliage ! reallo_frt reallocation parameter of fine root ! alphac ratio of coarse wood (twigs, branches, roots) to sapwood ! cr_frac fraction of tbc (twigs, branches, roots) that is coarse roots ! prhos* sapwood density [kg/m³} ! pnus foliage mass to sapwood area ratio [kg/cm**2] (pipe model) ! pha height growth parameter [cm/kg] ! pha_coeff1 height growth parameter coefficient 1 ! pha_coeff2 height growth parameter coefficient 2 ! pha_v1 height growth parameter 1 for non linear foliage height relationship ! pha_v2 height growth parameter 2 for non linear foliage height relationship ! pha_v3 height growth parameter 3 for non linear foliage height relationship ! crown_a parameter for crown radius - DBH relation [m/cm] ! crown_b parameter for crown radius - DBH relation [m] ! crown_c parameter for crown radius - DBH relation [m] ! psla_min* specific one-sided leaf area [m2/kg DW] ! psla_a ! phic* efficiency parameter, different for everg/decid [-] ! pnc N content (to be replaced by plant N model!) ! kco2_25 Michaelis constant of temperature (base 25 °C) [Pa] ! ko2_25 inhibition constant of O2 [kPa] (eqn 20) ! pc_25* CO2/O2 specificity value (base 25 °C) ! q10_kco2 Q10 values (acclimated to 25 °C ???) ! q10_ko2 ! q10_pc* ! pb Rd to Vm ratio !PItmin PIM: Inhibitor min temp. !PItopt PIM: Inhibitor opt temp. !PItmax PIM: Inhibitor max temp. !PIa PIM: Inhibitor scaling factor !PPtmin PIM: Promotor min temp. !PPtopt PIM: Promotor opt temp. !PPtmax PIM: Promotor max temp. !PPa PIM: Promotor scaling factor !PPb PIM: Promotor scaling factor !CSTbC CSM: chilling base temp. !CSTbT CSM: base temp. !CSa CSM: scaling factor !CSb CSM: scaling factor !LTbT TSM: base temp. !LTcrit TSM: critical temperature sum !Lstart TSM: start day after 1.11. !Phmodel used pheno model 0: no model, 1: PIM, 2: CSM, 3: TSM ! end_bb leaf shedding/ end of vegetation period ! fpar_mod modifying parameter in canopy_geometry ! ceppot_spec interception capacity parameter ! Nresp slope of photosynthesis response to N-limitation ! regflag flag for regeneration ! seedrate max. seeddling rate /m2 ! seedmass mass of a single seed [g] ! seedsd standard deviation of seed mass [g] ! seeda parameter of allometric relation shoot biomass - foliage bm ! seedb ---------"----------- ! pheight1 parameter of all. relation shoot biomass - height of seedling ! pheight2 ------------------" ----------- ! pheight3 ! pdiam1 parameter of all. relation shoot biomass - diameter at shoot basis ! pdiam2 ----------"----------------- ! pdiam3 ----------"-------------- ! k_opm_fol ! k_syn_fol ! k_opm_stem ! k_syn_stem ! k_opm_tb ! k_syn_tb ! k_opm_frt ! k_syn_frt ! k_opm_crt ! k_syn_crt ! nspecies 15 nspec_tree 13 !*************************************************************************************************************************************************************** beech 'Fagus sylvatica' spruce 'Picea abies' pine 'Pinus sylvestris' oak 'Quercus robur' birch 'Betula pendula' pinec 'Pinus contorta' pinep 'Pinus ponderosa' aspen 'Populus tremula' pineh 'Pinus halepensis' dougfir 'Pseudotsuga menziesii' robinia 'Robinia pseudoacacia' eglobulus 'Eucalyptus globulus' egrandis 'Eucalyptus grandis' grveg 'Ground vegetation' mistletoe 'Viscum alba' !*************************************************************************************************************************************************************** name 'beech' 'spruce' 'pine' 'oak' 'birch' 'pinec' 'pinep' 'aspen' 'pineh' 'dougfir' 'robinia' 'eglobulus' 'egrandis' 'grveg' 'mistletoe' amax 430 930 760 1060 220 600 850 160 250 930 200 50 50 5 30 yrec 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 stol 5 4 1 2 1 1 1 2 1 4 2 1 1 4 1 pfext 0.4 0.6 0.6 0.5 0.54 0.6 0.5 0.872 0.6 0.3 0.46 0.5 0.5 0.5 0.5 sigman 0.032 0.05 0.03 0.03 1.1 0.03 0.03 0.07508 0.03 0.025 0.176172 0.025 0.75 1 -99 respcoeff 0.5 0.52 0.52 0.5 0.5 0.71 0.44 0.5 0.52 0.5 0.5 0.53 0.53 0.5 0.5 prg 0.25 0.25 0.2 0.25 0.25 0.2 0.2 0.25 0.32 0.25 -99 0.25 0.25 0.25 0.25 prms 0.0035 0.00043 0.00024 0.0035 0.0035 0.000032 0.00024 0.0014 0.00024 -99 -99 0.3 0.3 0.0035 -99 prmr 0.0133 0.0033 0.007 0.01 0.01 0.00338 0.0007 0.0034 0.007 -99 0.1938 -99 -99 0.01 -99 psf 1 0.181 0.31 1 1 0.167 0.24 1 0.5 0.2 1 0.33 0.33 1 0.166 pss 0.026 0.022 0.04 0.05 0.02 0.023 0.05 0.04 0.04 0.05 0.25 0.03 0.03 0.4 -99 psr 0.65 0.5 0.5 0.5 0.5 0.4 0.8 0.5 1.095 0.75 0.67 1 1 0.75 -99 pcnr 0.008 0.0052 0.0079 0.008 0.02 0.0079 0.0212 0.008 0.0079 0.00955 0.0502 0.005 0.005 0.2 -99 ncon_fol 26.01 13.36 13.46 25 25 13.4 10.6 25 13.46 15.22 33.623 12 12 13.4 -99 ncon_frt 7.15 10.77 7.44 8.94 8.9 7.4 7.44 16 7.44 3.67 23.55 9.6 9.6 7.4 -99 ncon_crt 3.03 4.14 1.77 3.71 3.7 1.7 1.77 4 1.77 1.62 17.168 9.6 9.6 1.7 -99 ncon_tbc 4.27 5.24 3.61 6.19 5.4 3.6 3.61 6 3.61 3.62 17.16 3.8 3.8 3.6 -99 ncon_stem 1.54 1.22 1.09 2.1 1.7 1.1 1.09 0.6 1.09 1.035 15.345 1 1 1.1 -99 reallo_fol 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.6 0.1 0.1 0.1 0.5 0.5 0.1 0.1 reallo_frt 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.1 0.55 0.55 0.1 -99 alphac 0.48 0.5 0.46 0.56 0.5 0.24 0.46 0.46 0.5 0.54 0.51 0.22 0.22 0.5 -99 cr_frac 0.5 0.6 0.6 0.55 0.42 0.5 0.6 0.328 0.1 0.54 0.86 0.52 0.52 0.5 -99 prhos 0.00065 0.00042 0.000403 0.00056 0.0005365 0.00042 0.000434 0.000402 0.00062 0.000405 0.00078 0.0005 0.0005 -99 -99 pnus 0.03 0.096 0.05 0.02 0.025 0.046 0.022 0.02 0.043 0.093 0.05 0.04 0.06 0.03 -99 pha 125 40 190 100 150 190 190 150 190 40 -99 -99 -99 -99 -99 pha_coeff1 0.5 0.66666 0.66666 0.5 0.5 0.66666 0.66666 1.0823 0.66666 0.66666 -99 -99 -99 -99 -99 pha_coeff2 0.5 0.33333 0.33333 0.5 0.5 0.33333 0.33333 0.3189 0.33333 0.33333 -99 -99 -99 -99 -99 pha_v1 1089.3 284.8 206 946.7 900 1100 206 900 210 750 500 840 854 -99 -99 pha_v2 0.1351 -0.0151 0.03177 0.299 0.38 0.3 0.03177 0.08 0.08 -0.015 0.8541 -0.02973 -0.2 -99 -99 pha_v3 0.504 0.5039 0.877 0.948 0.9 0.4184 0.877 0.6 0.6 0.35 0.4404 0.291323 0.29277 -99 -99 crown_a 0.09571 0.06383 0.05213 0.095 0.0896 0.1437 0.05213 0.06 0.074 0.081287 0.0776 0.1249 0.1249 -99 -99 crown_b 0.57732 0.33567 0.48139 0.5 0.5716 0.1081 0.48139 0.4 0.255 0.355485 0.7368 0.7879 0.7879 -99 -99 crown_c 15 12 10 15 6 10 10 6 10 5 4 5.9 5.9 -99 -99 psla_min 12 3.78 4 14 10 7.5 3.1 7.2 4.84 2.82 13.2 15 0.18 20 4.18 psla_a 12 2.4 1 4.7 20 1.2 1 4.4 1 4.87 19.5 13.52 13.52 20 4.18 phic 1 0.8 0.9 1 1 0.9 0.9 1 0.9 0.8 1 1 1 1 0.8 pnc 20 20 20 20 17.9 50 20 24 20 45 33.623 -99 -99 10 -99 kco2_25 30 30 30 30 30 30 30 30 40.4 30 30 30 30 30 30 ko2_25 30 30 60 30 30 60 60 60 24.8 30 30 30 30 30 30 pc_25 3400 2600 3400 3400 3400 3400 3400 3400 3400 2600 3400 3400 3400 3400 3400 Q10_kco2 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 2.1 Q10_ko2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 1.2 Q10_pc 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 0.57 pb 0.01 0.015 0.01 0.01 0.035 0.01 0.01 0.0143 0.0241 0.015 0.01 0.035 0.035 0.01 0.015 PItmin -10.34 -99 -99 -23.05 -24.96 -99 -99 -10.34 -99 -99 -99 -99 -99 -99 -99 PItopt -0.89 -99 -99 -0.3 -10 -99 -99 -0.89 -99 -99 -99 -99 -99 -99 -99 PItmax 18.11 -99 -99 16.91 15.05 -99 -99 18.11 -99 -99 -99 -99 -99 -99 -99 PIa 0.058326 -99 -99 0.055149 0.030619 -99 -99 0.058326 -99 -99 -99 -99 -99 -99 -99 PPtmin -10.03 -99 -99 3.46 -7.03 -99 -99 -10.03 -99 -99 -99 -99 -99 -99 -99 PPtopt 28.61 -99 -99 34.55 21.8 -99 -99 28.61 -99 -99 -99 -99 -99 -99 -99 PPtmax 44.49 -99 -99 34.55 25.35 -99 -99 44.49 -99 -99 -99 -99 -99 -99 -99 PPa 0.109494 -99 -99 0.331253 0.064803 -99 -99 0.109494 -99 -99 -99 -99 -99 -99 -99 PPb 0.039178 -99 -99 0.010379 0.045432 -99 -99 0.039178 -99 -99 -99 -99 -99 -99 -99 CSTbC 11.03 -99 -99 18.31 18.42 -99 -99 11.03 -99 -99 -99 -99 -99 -99 -99 CSTbT 6.5 -99 -99 5.58 4.19 -99 -99 6.5 -99 -99 -99 -99 -99 -99 -99 CSa 3039.76 -99 -99 3451.18 3075.99 -99 -99 3039.76 -99 -99 -99 -99 -99 -99 -99 CSb -574.49 -99 -99 -631.5 -573.67 -99 -99 -574.49 -99 -99 -99 -99 -99 -99 -99 LTbT -6.98 -99 -99 0.49 -6.07 -99 -99 -6.98 -99 -99 11.5 -99 -99 -99 -99 LTcrit 664.88 -99 -99 372.49 672.9 -99 -99 664.88 -99 -99 350.9 -99 -99 -99 -99 Lstart 70 -99 -99 70 47 -99 -99 70 -99 -99 70 -99 -99 -99 -99 Phmodel 1 0 0 1 1 0 0 1 0 0 3 -99 -99 0 0 end_bb 282 366 366 287 278 366 366 282 366 366 282 366 366 302 366 fpar_mod 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ceppot_spe 0.6 0.8 0.9 0.5 0.5 0.4 0.4 0.3 0.9 0.8 0.5 0.3 0.3 0.5 0.6 Nresp 0.0068 0.0068 0.0062 0.0068 0.0068 0.0062 0.0062 0.0068 0.0062 0.0068 0.0068 0.0068 0.0068 0.006 0.0068 regflag 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 seedrate 1 1 1 1 1 1 1 1 1.5 1 1 1 1 -99 -99 seedmass 0.225 0.009 0.006 3.8 0.0002 0.005 0.0055 0.00016 0.019 0.01 0.0228 -99 -99 -99 -99 seedsd 0 0 0.0011268 0 0 0.0011268 0.0011268 0 0.0011268 0 0 -99 -99 -99 -99 seeda 0.398 0.8524 0.3927 0.2505 0.313 0.2609 0.3927 0.381 0.5044 1.3217 0.778 -99 -99 -99 -99 seedb 0.947 1.007 0.6786 0.7232 0.8 0.757 0.6786 0.8365 0.6486 1.1288 0.9645 -99 -99 -99 -99 pheight1 2.388 0.443 2.512 1.6947 4.651 1.5 2.512 0.2829 1.3145 5.7278 0.118272 -99 -99 -99 -99 pheight2 0.366 0.23 0.357 0.3896 0.3333 0.7236 0.357 0.57 0.3426 0.2443 0.757755 -99 -99 -99 -99 pheight3 0 0.0134 0 0 0 -99 0 -99 0 -99 -99 -99 -99 -99 -99 pdiam1 -1.637 -1.658 0 0 0 -99 0 -99 0 -99 -99 -99 -99 0.35 -99 pdiam2 0.433 0.386 0 0 0 -99 0 -99 0 -99 -99 -99 -99 0.35 -99 pdiam3 -0.0126 0 0 0 0 -99 0 -99 0 -99 -99 -99 -99 0.3 -99 k_opm_fol 0.02 0.06 0.05 0.015 0.01 0.005 0.015 0.04 0.002 0.08 0.3 0.5 0.5 0.03 0.01 k_syn_fol 0.3 0.2 0.1 0.4 0.78 0.6 0.8 0.4 0.5 0.2 0.2 0.01 0.01 0.2 0.5 k_opm_frt 0.02 0.05 0.05 0.01 0.00106 0.001 0.009 0.03 0.0004 0.05 0.008 0.001 0.001 0.001 -99 k_syn_frt 0.4 0.1 0.1 0.3 0.3 0.3 0.3 0.4 0.5 0.3 0.3 0.01 0.01 0.3 -99 k_opm_crt 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0.0009 0 -99 k_syn_crt 0.1 0.1 0 0.1 0.1 0.1 0 0.1 0 0.1 0.1 0.1 0.1 0.1 -99 k_opm_tb 0.006 0.006 0.006 0.006 0.0009 0.0009 0.0009 0.0009 0.006 0.006 0.0009 0.0009 0.0009 0 -99 k_syn_tb 0.5 0.8 0.5 0.5 0.8 0.5 0.5 0.5 0.5 0.8 0.8 0.8 0.8 0 -99 k_opm_stem 0.0025 0.0005 0.0005 0.0005 0.0005 0.0001 0.00055 0.0001 0.0005 0.0005 0.0005 0.0005 0.0005 0 -99 k_syn_stem 0.1 0.1 0 0.1 0.1 0.1 0 0.1 0 0.1 0.1 0.1 0.1 0 -99 spec_rl 30 15 15 30 30 15 15 30 15 15 30 30 30 60 -99 tbase 5 5 5 5 5 5 5 5 5 5 5 5 5 5 -99 topt 19.4 16.5 16.5 22 16.5 16.5 16.5 16.5 16.5 16.5 16.5 16.5 16.5 16.5 -99 bdmax_coef 1.55 1.62 1.62 1.55 1.55 1.62 1.62 1.55 1.62 1.62 1.55 1.55 1.55 1.62 -99 porcrit_co 0.4 0.7 0.7 0.4 0.4 0.7 0.7 0.4 0.7 0.7 0.4 0.4 0.4 0.7 -99 ph_opt_max 8.25 5.75 5.75 8.25 8.25 5.75 5.75 8.25 5.75 5.75 8.25 8.25 8.25 8.25 -99 ph_opt_min 4.25 3.75 3.75 4.25 3.75 3.75 3.75 4.25 3.75 3.75 3.75 3.75 3.75 3.5 -99 ph_max 9.25 7.75 7.75 9.25 9.25 7.75 7.75 9.25 7.75 7.75 9.25 9.25 9.25 9.25 -99 ph_min 3.75 2.25 2.25 3.75 2.25 2.25 2.25 3.75 2.25 2.25 2.25 2.25 2.25 2 -99 v_growth 1 1 1 1 1 1 1 1 1 1 1 1 1 5 -99