dynamic vegetation: yes - in terms of dimensional changes and natural mortality of trees. New individuals need to be introduced as a mangement event, there is no natural regeneration.
nitrogen limitation: yes - photosynthesis is reduced if nitrogen supply is not sufficient. Also, respiration increases with tissue nitrogen content and uptake.
CO2 effects: yes - photosynthesis is calculated in dependence on CO2 concentration (Farquhar approach)
light interception: yes - light distribution is calculated in canopy layers according to canopy properties. The model differentiates between direct and diffuse light fractions in each layer.
phenology: yes - the development of new leaves is driven by growing degree days (cumulative daily temperature above threshold). The demand for other supporting tissues (sapwood, fine roots) is defined due to their relationship to foliage and thus their growth follows that of foliage. Tissue senescence is empirically prescribed.
water stress: yes - photosynthesis is limited by stomatal conductance which in turn is decreased if water supply is not sufficient to fulfill the evaporative demand. In addition, there are some threshold values for foliage and fine root growth. Mortality is not affected by drought.
heat stress: no - (except that evaporative demand increases with temperature and stomata close with increasing vapor pressure deficit)
Evapo-transpiration approach: The model uses a modified Thornthwaite approach that calculates potential evapotranspiration based on temperature. The approach is modified to account for daily demand.
Root distribution over depth: Fine root distribution within the rooting depth is calculated empirically (species-specific).
closed energy balance: no (only latent heat, temperature and radiation input are calculated)
Coupling/feedback between soil moisture and surface temperature: yes - soil temperature is calculated in each soil layer according to the specific soil properties including water content.
latent heat: yes - total evapotranspiration is calculated as the sum of evaporation from interception, ground evaporation, and transpiration.
sensible heat: no